Along with the springing up of semantics-empowered communication research spanning from theories, applications, metrics to implementation, it is now witnessing an unprecedented growing interest in both academia and industry. In this work, we primarily aim to provide a comprehensive survey on both the background and research taxonomy, as well as a detailed technical tutorial for potential participants. Specifically, we start by reviewing the literature and answering the "what" and "why" questions in semantic transmissions. Afterwards, we present corresponding ecosystems, including theories, metrics, datasets and toolkits, on top of which the taxonomy for macro research directions is presented. Furthermore, we propose to categorize the critical enabling techniques by explicit and implicit reasoning-based methods, and elaborate on how they evolve and contribute to modern content & channel semantics-empowered communications. Besides reviewing and summarizing the latest efforts in semantics-empowered communications, we discuss the relations with other communication levels (e.g., reliable and goal-oriented communications) from a holistic and unified viewpoint. Subsequently, in order to facilitate the future developments and industrial applications, we also highlight advanced practical techniques for boosting semantic accuracy, robustness, and large-scale scalability, just to mention a few. Finally, we discuss the technical challenges that shed light on future research opportunities. Considering the growing yet the ever-updating status of semantics-empowered communications, we hope this work would benefit not only sophisticated but also intended practitioners.
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